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a collection of species that occur together in the same habitat at one time

definition of community

a group of species with similar lifestyles, epiphytes

definition of guild, provide and example

emergent, canopy, understorey, shrub layer, forest floor

tiers in a forest (5)


example of an emergent species in New Zealand


example of a canopy species in New Zealand


example of an understorey species in New Zealand


the struggle between organisms to survive in a habitat with limited resources; stronger within a guild, strongest within a species (intraspecific).


interaction in which one organism captures and feeds on another organism; a key problem for sessile plants which is accommodated through mechanical and chemical defenses.


form of predation that leaves the host alive (e.g. mistletoe)


one species is neutrally affected and one species benefits (e.g. epiphytes)


both species benefit (e.g. mychorrhizae on plant roots)


*mutualistic interaction in which a fungus grows on plant root hairs, increasing the root surface area so plants experience increased nutrient uptake and the fungus leeches food from the plant.


spatial change is caused by changes in __________________


large scale spatial change occurs between ____________________

lowland tropical rainforest

biome at equatorial latitudes, dominated by angiosperms, high spp. diversity, spp. with tolerance of higher temperatures

warm temperate rainforest

biome at mid latitudes, dominated by angiosperms and gymnosperms, moderate spp. diversity, app. with tolerance of cooler temperatures (e.g. winter deciduous)

boreal forest

biome at high latitudes, dominated by gymnosperms, low spp. diversity, app. tolerant of low temperatures


biome at polar latitudes, very few spp., low-growing plants adapted to cold/wet conditions

vegetative zonation, elevation

_____________________ is an example of a moderate scale spatial change, and is caused by changes in _____________________.


tree with a broad elevation tolerance (little sensitivity to elevation gradient)


tree with a narrow elevation tolerance (high elevations)

forest zonation, flood frequency

_____________________ is an example of a small scale spatial change, and is caused by changes in _____________________.


nutrient demanding tree that requires flooding every few years


dry-foot tree, has lower nutrient needs and requires decadal flooding


wet-foot tree, huge biomass and thus huge nutrient demands, requires annual flooding


scientist who pioneered the field of plant succession

directional series of changes over time in the environmental condition and species composition at a particular site


pioneer, mid-successional, mature phase

three stages in primary succession

access to light

principle changing variable in succession


succession is niche defined in ________________.

mature phase forest

late successional forest that experiences low light and intense competition


after a _________________ (e.g. volcanic erruption, tree fall gap), the site experiences high light, forest regeneration; promotes spp. diversity

kanuka, manuka

two examples of pioneer guild plants

pioneer guild

guild that lasts 50-80 years, examples of which include manuka and kanuka

kanuka, light-demanding, stress-tolerant, highly dispersive, a weak competitor, disturbance-dependent, short-lived

a pioneer guild species like _______________ is generally (6):

mamangi, maire

two examples mid-successional guild plants

kohekohe, taraire, kauri

three examples late-successional guild plants

kohekohe, shade-tolerant, stress-intolerant, less dispersive, a strong competitor, disturbance independent, long-lived

a late-successional guild species like _______________ is generally (6):

riparian zonation, exposure and substrate

_____________________ is an example of a very small scale spatial change, and is caused by changes in _____________________.


gymnosperm example of a Gondwanan plant


angiosperm example of a Gondwanan plant

Horopito, plesiomorphic

an angiosperm example of a Gondwanan that has a basal or ________________ flower with leaf-like stamens and open carpels without styles


_________________ are also evidence of Gondwanan origins because they occur in Aus./NZ, S. America, and New Caledonia, but their seeds can't disperse further than across 25 km of sea


_________________ provide evidence of past vegetation because they are sites of pollen deposition and preservation.

instantaneous, 400

the transition from glacial cold-tolerant community to a warm climate forest was almost ____________________; it lasted around ____________________ years.


the first colonizer in post-glacial recovery


the second colonizer in post-glacial recovery

Southern Rata

part of the myrtle family, this tree is was dispersed from Australia/Antarctica (non-Gondwanan)


a plant of non-Gondwanan origins that's part of the mahogany family

tree-ferns, Nothofagus

during the Holocene era, initial vegetative dominance was by dispersive __________________ in montane habitats, followed by the slow-spreading of ____________________.

pleistocene, grass-shrubland, Northland/Coromandel, shrubland, tussock-herbfield

during the ___________________ epoch, New Zealand experienced periods of glaciation which became increasingly more severe. _____________________ was widespread, forests were dominant only in _____________________. In the North Island, subalpine _____________________ was dominant south of Auckland, and in the South Island, _____________________ was widespread.

frugivory, attachment, flotation, wind

modes of plant dispersal in order of frequency (4)

podocarps, gymnosperm, 60

_____________________ have _____________________ leaves: they're small and scale-like, have parallel venation, and have __% of the photosynthetic efficiency of broadleaf Angiosperms.

light-dependent, disturbance-dependent, long-lived

podocarp seedlings are (3):

podocarps, shade-tolerant, light-demanding

Rimu, Kahikatea, Totara, Matai, and Miro are all examples of _________________. Miro is an exception because it's _____________________, whereas the first four are _____________________.

broadleaf angiosperms

Puriri, Rewarewa, Kohekohe, Mangeao, Tawa, Kamahi, Manuka, and Kanuka are all examples of:


_____________________ pioneer small gaps.

Lake Taupo, Nuee Ardente, podocarps

_____________________ experienced _____________________ volcanic flow, giving the stress-tolerant _____________________ the advantage.

dense podocarp forest, emergents, Angiosperms

Whirinaki is a _____________________, unusually dominated by _____________________; _____________________ form the main canopy, trees are present but suppressed.

manuka, kanuka, podocarp, podocarp, natural thinning, podocarps, species composition

Large scale regeneration cycle: 1, _____________________/_____________________ scrub, abundant _____________________ seedlings; 2, _____________________ pole stage, stand depresses kanuka etc.; 3, _____________________ of _____________________ as they increase in height; 4, shift in _____________________

tree ferns, kamahi, tree ferns, Tawa, podocarp, podocarps

Gap regeneration cycle: 1, gap created by fall of large tree is colonized by _____________________; 2, _____________________ grows and suppresses the _____________________, _____________________ on gap margins, _____________________ seedlings beneath; 3, rapid growth of pole _____________________ to occupy gap

podocarps, hardwoods, beeches

As altitude increases, vegetation zonation is _____________________ -> _____________________ -> _____________________

pumice, broadleafs, podocarps

_____________________ substrate holds more water, thus producing more stressful conditions that _____________________ can't tolerate, but _____________________ can

Rimu, semi-shade-tolerant, mixed

*(relatively) _____________________ podocarp, _____________________-age stand, medium colonization length

Miro, most shade-tolerant, mixed

*(relatively) _____________________ podocarp, _____________________-age stand, longest colonization length, ongoing juvenile recruitment

Matai, light-demanding, even

*(relatively) _____________________ podocarp, _____________________-age stand, one cohort, shortest colonization length

Tawa, Kohekohe, gap-dependent, gaps, frost

*two strongly shade-tolerant broadleafs in Rotoehu Forest; not _____________________ for regeneration but rather negatively affected by __________________ due to ____________________ sensitivity.

Mangeao, Rewarewa, small gap, all

*two more light-demanding broadleafs in Rotoehu Forest; _____________________ specialists, seedlings of ______________ size class(es)

Tawa, Kamahi, Rimu, Miro, Tawa

*_____________________ dominant in Urewera National park, _____________________, _____________________, and _____________________ suppressed; fewer _____________________ at higher elevations, others increase in abundance

archaic conifer

being from 200 million years ago, Kauri are an example of an _____________________.

ecosystem engineers

Kauri are _____________________, creating alternate stable states.

alternate stable state

in presence of plant, like Kauri, the community is differently structured than in absence

Kohekohe, puriri

two main competitors of Kauri

Toru, Tawari

*two kauri associate species that experience depressed nutrient cycling

slow organic decay, litter, cup podsol, depressed

Kauri as an ecosystem engineer: _____________________ under Kauri causing the creation of a deep layer of _____________________; creation of a _____________________ underneath tree; _____________________ cycling

terpenes, waxes, phenols, inhibit bacteria and fungi

three methods to slow decay of Kauri litter and what these four methods do

broadleaf, conifer

_____________________ litter is thinner wheras _____________________ litter is generally thicker

cup podsol, acidic, neutralizes nutrients, leaching

*Kauri creates a _____________________: litter is _____________________, which _____________________ and flushes them deeper, promoting _____________________

nitrogen, ammonium, nitrate

Kauri depressed nutrient cycling converts _____________________ to _____________________ which is toxic for other plants, rather than _____________________ which is generally better for plants

ridge crest specialists, rapid podsolization, high, frequent, long-lived

*Kauri are _____________________ which helps with _____________________ and access to _____________________ light, _____________________ disturbance. They are _____________________-lived

short recruitment, competition, self-thinning, competition, recruitment, synchronized disturbance, biomass, broadleafs, ridgecrest

1st Kauri cohort after disturbance: _____________________ phase due to lack of _____________________, followed by a _____________________ and mature phase due to intense _____________________. 2nd cohort: longer _____________________ phase without _____________________, part of 2nd cohort _____________________ replaced by _____________________ as Kauri retreat to _____________________.

retreated north, returned south, mean daily maximum temperature

*Kauri _____________________ during glaciation and _____________________ post-glaciation, because they require a _____________________ of 17 oC

Matai, softwood, self-preserved

_____________________ is a _____________________ that produces superior timber, a tall tree that's straight grained. Its timber is _____________________ and richly colored

Tawa, hardwood, decays faster, Red Beech, hardwood

_____________________ is a _____________________ that produces an inferior timber, a smaller log which _____________________. _____________________ is another example of a _____________________.

Nothofagus, erratic, best, collapsing residual

in north Westland, a remnant _____________________ forest, logging was _____________________ and piecemeal, targeting only the _____________________ trees, leaving a _____________________ stand.

decreased, increase, remained constant, increased, remained constant, disease, insect attack, pinhole borer

Managed beach forest dynamics: density _____________________, residual diameter _____________________, basal area _____________________, height and volume _____________________. After 100 years: volume _____________________, and stems are large enough to harvest and should be, rather than waiting for _____________________ or _____________________ (e.g. _____________________)

aggressive, more careful, Tawa, too small

*Tihoi: _____________________ harvest; Whirinaki: _____________________ harvest, improved stand stability, but regeneration by _____________________ because gaps created were _____________________ for podocarp regeneration.

coup felling, podocarps, large disturbance, stand senesence

*Saltwater Forest: Wetland silviculture model: _____________________, advantages valuable _____________________, mimics _____________________ regime, harvest just as _____________________ begins

Kauri, ridge crests, unfamiliar slope position, ringbarking, expensive, time-consuming

_____________________ is the most valuable timber, attempted planting all over slopes (not just on _____________________), poor survival of planted juveniles in _____________________, _____________________ of competitors. this method is too _____________________ and _____________________, and therefore not feasible.

recreational, conservation

the modern view of silviculture holds that forests have greater _____________________ and _____________________ blue than they do as timber

Hard Beech

most northerly, most lowland, often found in forest with kauri, abundant in lower montane of northern Southland, occurs in ecotonal forest with lowland species

Red Beech

largest, abundant in lower montane throughout Southland and Bay of Plenty, like hard beech, occurs in ecotonal forest with lowland species, can be found in mixed Podocarp forests

Black Beech

often grows with Mountain Beech, found in eastern NZ in foothills at ridge crest (where it's drier), poor soil specialists, mixed with lowland spp. in lower montane

Mountain Beech

often grows with Black Beech, high elevation, lower to upper montane, found in drier east central North Island southwards

Silver Beech

found in wetter west, Bay of Plenty southwards, high rainfall, lower to upper montane, forming treeline

shade-tolerant, mixed

Silver beeches are _______________ (light-damanding/shade-tolerant) and structured in __________ (mixed/even) -age stands.


Beech forest tree species diversity _____________ as altitude (and stress) increases, but not herb and shrub spp. diversity


*Beech forest spp. diversity ______________ as rainfall decreases

light-demanding, even

Black beeches are _______________ (light-damanding/shade-tolerant) and structured in __________ (mixed/even) -age stands.

light-demanding, even

Mountain beeches are _______________ (light-damanding/shade-tolerant) and structured in __________ (mixed/even) -age stands.

light-demanding, even

Hard beeches are _______________ (light-damanding/shade-tolerant) and structured in __________ (mixed/even) -age stands.

light-demanding, even

Red beeches are _______________ (light-damanding/shade-tolerant) and structured in __________ (mixed/even) -age stands.

increase, level out

*during periods of stand development, biomass tends to first ______________ and then __________________ as the stand reaches maturity.

even-aged in distinct groups

*in Saltwater Forest, the stand structure that facilitated coup-felling was:

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